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The Origin of Species · Chapter

I: Variation Under Domestication explained

Darwin introduces the concept of variation under domestication, arguing that domestic organisms vary more than wild ones due to changed conditions of life.

What happens

Darwin introduces the concept of variation under domestication, arguing that domestic organisms vary more than wild ones due to changed conditions of life. He discusses causes of variability, the roles of definite and indefinite variation, and the importance of selection by man in shaping domestic breeds. He also addresses the difficulty of distinguishing varieties from species and the origin of domestic races from one or more wild stocks.

Themes in this chapter

  • Domestication and Artificial Selection

    Central theme: Darwin explains how human selection drives variation and the formation of domestic races.

  • Variation and Inheritance

    Explored through causes of variability, definite and indefinite variation, and inheritance of traits under domestication.

Characters to notice

  • Charles Darwin

    Author and narrator, presenting his observations and conclusions on variation under domestication.

  • Professor Weismann

    Cited for his insistence on the two factors of variation: the nature of the organism and the nature of the conditions.

  • Kölreuter

    Mentioned in the context of the reproductive system's sensitivity to changed conditions and variability.

  • Mr. Youatt

    Referenced for his emphasis on selection as the means of improving domestic breeds.

Key passages

  • When we compare the individuals of the same variety or sub-variety of our older cultivated plants and animals, one of the first points which strikes us is, that they generally differ more from each other than do the individuals of any one species or variety in a state of nature.

    Domesticated plants and animals show greater individual variation than their wild counterparts.

    Darwin's opening observation on variability under domestication.

  • Man can hardly select, or only with much difficulty, any deviation of structure excepting such as is externally visible; and indeed he rarely cares for what is internal.

    Human selection focuses on external traits, not internal anatomy.

    Explains why domestic breeds often differ greatly in external features but less in internal organs.